Thursday, August 21, 2008

Photosynthesis and oxygen

Photosynthesis and oxygen

The harnessing of the sun’s energy led to several major changes in life on Earth.
The harnessing of the sun’s energy led to several major changes in life on Earth.

It is likely that the initial cells were all heterotrophs, using surrounding organic molecules (including those from other cells) as raw material and an energy source. As the food supply diminished, a new strategy evolved in some cells. Instead of relying on the diminishing amounts of free-existing organic molecules, these cells adopted sunlight as an energy source. Estimates vary, but by about 3 billion years ago, something similar to modern photosynthesis had probably developed. This made the sun’s energy available not only to autotrophs but also to the heterotrophs that consumed them. Photosynthesis used the plentiful carbon dioxide and water as raw materials and, with the energy of sunlight, produced energy-rich organic molecules (carbohydrates).

Moreover, oxygen was produced as a waste product of photosynthesis. At first it became bound up with limestone, iron, and other minerals. There is substantial proof of this in iron-oxide rich layers in geological strata that correspond with this time period. The oceans would have turned to a green color while oxygen was reacting with minerals. When the reactions stopped, oxygen could finally enter the atmosphere. Though each cell only produced a minute amount of oxygen, the combined metabolism of many cells over a vast period of time transformed Earth’s atmosphere to its current state. Among the oldest examples of oxygen-producing lifeforms are fossil Stromatolites.

This, then, is Earth’s third atmosphere. Some of the oxygen was stimulated by incoming ultraviolet radiation to form ozone, which collected in a layer near the upper part of the atmosphere. The ozone layer absorbed, and still absorbs, a significant amount of the ultraviolet radiation that once had passed through the atmosphere. It allowed cells to colonize the surface of the ocean and ultimately the land; without the ozone layer, ultraviolet radiation bombarding the surface would have caused unsustainable levels of mutation in exposed cells. Besides making large amounts of energy available to life-forms and blocking ultraviolet radiation, the effects of photosynthesis had a third, major, and world-changing impact. Oxygen was toxic; probably much life on Earth died out as its levels rose (the “Oxygen Catastrophe”).Resistant forms survived and thrived, and some developed the ability to use oxygen to enhance their metabolism and derive more energy from the same food.

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